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@ARTICLE{Delaforge:626391,
author = {Delaforge, Elise and Due, Amanda D and Theisen, Frederik
Friis and Morffy, Nicolas and O’Shea, Charlotte and
Blackledge, Martin and Strader, Lucia C and Skriver, Karen
and Kragelund, Birthe B},
title = {{A}llovalent scavenging of activation domains in the
transcription factor {ANAC}013 gears transcriptional
regulation},
journal = {Nucleic acids symposium series},
volume = {53},
number = {4},
issn = {0305-1048},
address = {Oxford},
publisher = {Oxford Univ. Press},
reportid = {PUBDB-2025-01383},
pages = {gkaf065},
year = {2025},
note = {ISSN 1362-4962 not unique: **2 hits**.},
abstract = {Transcriptional regulation involves interactions between
transcription factors, coregulators, and DNA. Intrinsic
disorder is a major player in this regulation, but
mechanisms driven by disorder remain elusive. Here, we
address molecular communication within the stress-regulating
Arabidopsis thaliana transcription factor ANAC013. Through
high-throughput screening of ANAC013 for transcriptional
activation activity, we identify three activation domains
within its C-terminal intrinsically disordered region. Two
of these overlap with acidic islands and form dynamic
interactions with the DNA-binding domain and are released,
not only upon binding of target promoter DNA, but also by
nonspecific DNA. We show that independently of DNA binding,
the RST (RCD--SRO--TAF4) domain of the negative regulator
RCD1 (Radical-induced Cell Death1) scavenges the two acidic
activation domains positioned vis-à-vis through allovalent
binding, leading to dynamic occupation at enhanced affinity.
We propose an allovalency model for transcriptional
regulation, where sequentially close activation domains in
both DNA-bound and DNA-free states allow for efficient
regulation. The model is likely relevant for many
transcription factor systems, explaining the functional
advantage of carrying sequentially close activation
domains.},
cin = {EMBL-User},
ddc = {540},
cid = {I:(DE-H253)EMBL-User-20120814},
pnm = {6G3 - PETRA III (DESY) (POF4-6G3)},
pid = {G:(DE-HGF)POF4-6G3},
experiment = {EXP:(DE-H253)P-P12-20150101},
typ = {PUB:(DE-HGF)16},
UT = {WOS:001417619200001},
doi = {10.1093/nar/gkaf065},
url = {https://bib-pubdb1.desy.de/record/626391},
}