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@ARTICLE{Boudinot:639693,
author = {Boudinot, Brendon and Beutel, Rolf G. and Weingardt,
Michael and van de Kamp, Thomas and Hammel, Jörg U. and Li,
Di and Richter, Adrian and Wipfler, Benjamin},
title = {{T}he insect head rewound: {C}larifications to the
groundplan of {H}exapoda ({P}ancrustacea)},
journal = {Arthropod structure $\&$ development},
volume = {89},
issn = {1467-8039},
address = {Amsterdam [u.a.]},
publisher = {Elsevier Science},
reportid = {PUBDB-2025-04620},
pages = {101490},
year = {2025},
abstract = {The organization of the hexapod head remains a cornerstone
problem in arthropod systematics, central to segmental
homology, character definition, higher-level phylogeny, and
functional and evolutionary morphology. Recently, Nel et al.
(2025) proposed an alternative interpretation of insect head
segmentation that departs markedly from established
anatomical and comparative frameworks. We evaluate the
internal consistency and external coherence of that
hypothesis using their model groups, broader taxonomic
samples, prior studies (including crustaceomorph
Pancrustacea) and our own investigations. We identify
multiple implausible anatomical interpretations and logical
contradictions in their reconstruction. Our reanalysis,
supported by microtomographic imaging (μ- and SR-μ-CT),
shows that the proposed revision lacks empirical foundation
(e.g. Psocodea), does not meet its own definitions (e.g.
Neuropteroidea), misidentifies homologs (e.g. Coleoptera,
Formicidae), and is unsupported by fossil or developmental
evidence. Consequently, the new theory yields unreliable
homology statements and obscures groundplan conditions and
character polarities of the hexapod head. We therefore
reject the hypothesis that “intercalate” and
“promandible” sclerites existed in the groundplan of
Hexapoda, and the assumed plesiomorphy of dicondyly. Our
findings clarify relationships among major head sclerites,
endoskeletal elements, and the head capsule's strengthening
ridges, underscoring the need for comprehensive anatomy,
broad sampling, and logical rigor in resolving arthropod
head evolution.},
cin = {DOOR ; HAS-User / Hereon},
ddc = {590},
cid = {I:(DE-H253)HAS-User-20120731 / I:(DE-H253)Hereon-20210428},
pnm = {6G3 - PETRA III (DESY) (POF4-6G3)},
pid = {G:(DE-HGF)POF4-6G3},
experiment = {EXP:(DE-H253)P-P05-20150101},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:41160967},
doi = {10.1016/j.asd.2025.101490},
url = {https://bib-pubdb1.desy.de/record/639693},
}