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@ARTICLE{Vilhelmsen:623159,
author = {Vilhelmsen, Lars and Boudinot, Brendon E. and Hammel, Jörg
U. and Nakamine, Hiroshi and Yamamoto, Shûhei},
title = {{A}n insect † {A}rchaeopteryx : {C}retaceous amber fossil
elucidates the evolution of complex host detection and
ovipositor mechanisms in parasitoid woodwasps
({H}ymenoptera: {O}russoidea)},
journal = {Systematic entomology},
volume = {50},
number = {3},
issn = {0307-6970},
address = {Oxford [u.a.]},
publisher = {Wiley-Blackwell},
reportid = {PUBDB-2025-00625},
pages = {630 - 645},
year = {2025},
abstract = {We describe †Cretovelona orussopteryx n. gen. $\&$ sp., a
female orussoid from mid-Cretaceous Kachin amber. We examine
the fossil with synchrotron scanning and integrate it into
an existing morphological data set for Orussoidea.
†Cretovelona is placed as sister to crown group Orussoidea
by Bayesian phylogenetic analyses. It is unique in
displaying a combination of plesiomorphic (e.g. 12
antennomeres, exposed ovipositor sheaths, partly
internalized ovipositor that does not extend further than
anterior part of abdomen) and apomorphic characters (e.g.
antenna with club, fore tarsus 3-segmented) that display
intermediate stages in the evolution of the host detection
mechanism and the fully internalized ovipositor in extant
Orussidae. The wing venation of †Cretovelona is even more
reduced than observed in extant Orussidae, likely correlated
with the small body size (<2 mm). Tracing the changes in
body size across the orussoid tree indicates that the stem
group underwent a size reduction event in the Cretaceous,
something that might have influenced character evolution in
for example wing venation and internalization of the
ovipositor. †Cretovelona is a rare example of a
transitional fossil that elucidates the evolution of complex
features when adapting to a highly specialized lifestyle, in
this case detecting and targeting hosts deep inside wood. We
propose to recognize only the family Orussidae within
Orussoidea. Orussidae comprises the crown group Orussoidea,
including all Cenozoic fossils described so far. The stem
group taxa, all Mesozoic fossils, are not assigned to any
specific family within Orussoidea. Zoobank Registration:
urn:lsid:zoobank.org:pub:397F76A5-7467-4BCA-9963-6A380EC05200.},
cin = {DOOR ; HAS-User / Hereon},
ddc = {570},
cid = {I:(DE-H253)HAS-User-20120731 / I:(DE-H253)Hereon-20210428},
pnm = {6G3 - PETRA III (DESY) (POF4-6G3) / FS-Proposal:
BAG-20230681 (BAG-20230681)},
pid = {G:(DE-HGF)POF4-6G3 / G:(DE-H253)BAG-20230681},
experiment = {EXP:(DE-H253)P-P05-20150101},
typ = {PUB:(DE-HGF)16},
UT = {WOS:001410844900001},
doi = {10.1111/syen.12673},
url = {https://bib-pubdb1.desy.de/record/623159},
}