TY - JOUR AU - Jaciuk, Marcin AU - Scherf, David AU - Kaszuba, Karol AU - Gaik, Monika AU - Rau, Alexander AU - Kościelniak, Anna AU - Krutyhołowa, Rościsław AU - Rawski, Michał AU - Indyka, Paulina AU - Graziadei, Andrea AU - Chramiec-Głąbik, Andrzej AU - Biela, Anna AU - Dobosz, Dominika AU - Lin, Ting-Yu AU - Abbassi, Nour-el-Hana AU - Hammermeister, Alexander AU - Rappsilber, Juri AU - Kosinski, Jan AU - Schaffrath, Raffael AU - Glatt, Sebastian TI - Cryo-EM structure of the fully assembled Elongator complex JO - Nucleic acids symposium series VL - 51 IS - 5 SN - 0305-1048 CY - Oxford PB - Oxford Univ. Press M1 - PUBDB-2024-00571 SP - 2011 - 2032 PY - 2023 N1 - ISSN 1362-4962 not unique: **2 hits**. AB - Transfer RNA (tRNA) molecules are essential to decode messenger RNA codons during protein synthesis. All known tRNAs are heavily modified at multiple positions through post-transcriptional addition of chemical groups. Modifications in the tRNA anticodons are directly influencing ribosome decoding and dynamics during translation elongation and are crucial for maintaining proteome integrity. In eukaryotes, wobble uridines are modified by Elongator, a large and highly conserved macromolecular complex. Elongator consists of two subcomplexes, namely Elp123 containing the enzymatically active Elp3 subunit and the associated Elp456 hetero-hexamer. The structure of the fully assembled complex and the function of the Elp456 subcomplex have remained elusive. Here, we show the cryo-electron microscopy structure of yeast Elongator at an overall resolution of 4.3 Å. We validate the obtained structure by complementary mutational analyses in vitro and in vivo. In addition, we determined various structures of the murine Elongator complex, including the fully assembled mouse Elongator complex at 5.9 Å resolution. Our results confirm the structural conservation of Elongator and its intermediates among eukaryotes. Furthermore, we complement our analyses with the biochemical characterization of the assembled human Elongator. Our results provide the molecular basis for the assembly of Elongator and its tRNA modification activity in eukaryotes. LB - PUB:(DE-HGF)16 C6 - pmid:36617428 UR - <Go to ISI:>//WOS:000912446800001 DO - DOI:10.1093/nar/gkac1232 UR - https://bib-pubdb1.desy.de/record/602296 ER -