TY - JOUR AU - Lewis, Hannah C. AU - Kelnhofer-Millevolte, Laurel E. AU - Brinkley, Mia R. AU - Arbach, Hannah E. AU - Arnold, Edward A. AU - Sanders, Saskia AU - Bosse, Jens Bernhard AU - Ramachandran, Srinivas AU - Avgousti, Daphne C. TI - HSV-1 exploits host heterochromatin for nuclear egress JO - The journal of cell biology VL - 222 IS - 9 SN - 0021-9525 CY - New York, NY PB - Rockefeller Univ. Press M1 - PUBDB-2024-00335 SP - e202304106 PY - 2023 AB - Herpes simplex virus (HSV-1) progeny form in the nucleus and exit to successfully infect other cells. Newly formed capsids navigate complex chromatin architecture to reach the inner nuclear membrane (INM) and egress. Here, we demonstrate by transmission electron microscopy (TEM) that HSV-1 capsids traverse heterochromatin associated with trimethylation on histone H3 lysine 27 (H3K27me3) and the histone variant macroH2A1. Through chromatin profiling during infection, we revealed global redistribution of these marks whereby massive host genomic regions bound by macroH2A1 and H3K27me3 correlate with decreased host transcription in active compartments. We found that the loss of these markers resulted in significantly lower viral titers but did not impact viral genome or protein accumulation. Strikingly, we discovered that loss of macroH2A1 or H3K27me3 resulted in nuclear trapping of capsids. Finally, by live-capsid tracking, we quantified this decreased capsid movement. Thus, our work demonstrates that HSV-1 takes advantage of the dynamic nature of host heterochromatin formation during infection for efficient nuclear egress. LB - PUB:(DE-HGF)16 C6 - pmid:37516914 UR - <Go to ISI:>//WOS:001068239900001 DO - DOI:10.1083/jcb.202304106 UR - https://bib-pubdb1.desy.de/record/601642 ER -