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@ARTICLE{Dubey:570307,
author = {Dubey, Badri Nath and Shyp, Viktoriya and Fucile, Geoffrey
and Sondermann, Holger and Jenal, Urs and Schirmer, Tilman},
title = {{M}utant structure of metabolic switch protein in complex
with monomeric c-di-{GMP} reveals a potential mechanism of
protein mediated ligand dimerization},
journal = {Scientific reports},
volume = {13},
number = {1},
issn = {2045-2322},
address = {[London]},
publisher = {Macmillan Publishers Limited, part of Springer Nature},
reportid = {PUBDB-2023-00670},
pages = {2727},
year = {2023},
abstract = {Bacterial second messengers c-di-GMP and (p)ppGpp have
broad functional repertoires ranging from growth and cell
cycle control to the regulation of biofilm formation and
virulence. The recent identification of SmbA, an effector
protein from Caulobacter crescentus that is jointly targeted
by both signaling molecules, has opened up studies on how
these global bacterial networks interact. C-di-GMP and
(p)ppGpp compete for the same SmbA binding site, with a
dimer of the c-di-GMP inducing a conformational change that
involves loop 7 of the protein thats leads to downstream
signaling. Here, we report a crystal structure of a partial
loop 7 deletion mutant, SmbA∆loop in complex with c-di-GMP
determined at 1.4 Å resolution. SmbA∆loop binds monomeric
c-di-GMP indicating that loop 7 is required for c-di-GMP
dimerization. Thus the complex probably represents the first
step of consecutive c-di-GMP binding to form intercalated
dimer as has been observed in wild-type SmbA. . Considering
the prevalence of intercalated c-di-GMP molecules observed
bound to proteins, the proposed mechanism may be generally
applicable to protein-mediated c-di-GMP dimerization.
Notably, in the crystal, SmbA∆loop forms a 2-fold
symmetric dimer via isologous interactions with the two
symmetric halves of c-di-GMP. Structural comparisons of
SmbA∆loop with wild-type SmbA in complex with dimeric
c-di-GMP or ppGpp support the idea that loop 7 is critical
for SmbA function by interacting with downstream partners.
Our results also underscore the flexibility of c-di-GMP, to
allow binding to the symmetric SmbA∆loop dimer interface.
It is envisaged that such isologous interactions of c-di-GMP
could be observed in hitherto unrecognized targets.},
cin = {CSSB-DESY-HS},
ddc = {600},
cid = {I:(DE-H253)CSSB-DESY-HS-20210521},
pnm = {633 - Life Sciences – Building Blocks of Life: Structure
and Function (POF4-633) / 6G3 - PETRA III (DESY) (POF4-6G3)},
pid = {G:(DE-HGF)POF4-633 / G:(DE-HGF)POF4-6G3},
experiment = {EXP:(DE-H253)P-P03-20150101},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:36810577},
UT = {WOS:001003439800022},
doi = {10.1038/s41598-023-29110-0},
url = {https://bib-pubdb1.desy.de/record/570307},
}