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@ARTICLE{Rouillon:491620,
author = {Rouillon, Christophe and Schneberger, Niels and Chi,
Haotian and Blumenstock, Katja and Da Vela, Stefano and
Ackermann, Katrin and Moecking, Jonas and Peter, Martin and
Boenigk, Wolfgang and Seifert, Reinhard and Bode, Bela E.
and Schmid-Burgk, Jonathan L. and Svergun, Dmitri and Geyer,
Matthias and White, Malcolm F. and Hagelueken, Gregor},
title = {{A}ntiviral signalling by a cyclic nucleotide activated
{CRISPR} protease},
journal = {Nature},
volume = {614},
number = {7946},
issn = {0028-0836},
address = {London [u.a.]},
publisher = {Nature Publ. Group},
reportid = {PUBDB-2023-00312},
pages = {168 - 174},
year = {2022},
abstract = {CRISPR defence systems such as the well-known DNA-targeting
Cas9 and the RNA-targeting type III systems are widespread
in prokaryotes1,2. The latter orchestrates a complex
antiviral response that is initiated through the synthesis
of cyclic oligoadenylates after recognition of foreign
RNA3,4,5. Among the large set of proteins that are linked to
type III systems and predicted to bind cyclic
oligoadenylates6,7, a CRISPR-associated Lon protease (CalpL)
stood out to us. CalpL contains a sensor domain of the SAVED
family7 fused to a Lon protease effector domain. However,
the mode of action of this effector is unknown. Here we
report the structure and function of CalpL and show that
this soluble protein forms a stable tripartite complex with
two other proteins, CalpT and CalpS, that are encoded on the
same operon. After activation by cyclic tetra-adenylate
(cA4), CalpL oligomerizes and specifically cleaves the MazF
homologue CalpT, which releases the extracytoplasmic
function σ factor CalpS from the complex. Our data
provide a direct connection between CRISPR-based detection
of foreign nucleic acids and transcriptional regulation.
Furthermore, the presence of a SAVED domain that binds
cyclic tetra-adenylate in a CRISPR effector reveals a link
to the cyclic-oligonucleotide-based antiphage signalling
system.},
cin = {EMBL-User / EMBL},
ddc = {500},
cid = {I:(DE-H253)EMBL-User-20120814 / I:(DE-H253)EMBL-20120731},
pnm = {6G3 - PETRA III (DESY) (POF4-6G3) / DFG project
G:(GEPRIS)390873048 - EXC 2151: ImmunoSensation2 - the
immune sensory system (390873048)},
pid = {G:(DE-HGF)POF4-6G3 / G:(GEPRIS)390873048},
experiment = {EXP:(DE-H253)P-P12-20150101 / EXP:(DE-H253)P-P13-20150101},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:36423657},
UT = {WOS:000912338800001},
doi = {10.1038/s41586-022-05571-7},
url = {https://bib-pubdb1.desy.de/record/491620},
}