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@ARTICLE{Pazicky:483482,
author = {Pazicky, Samuel and Werle, Anna-Leoni A. and Lei, Jian and
Loew, Christian and Weininger, Ulrich},
title = {{I}mpact of distant peptide substrate residues on enzymatic
activity of {S}ly{D}},
journal = {Cellular and molecular life sciences},
volume = {79},
number = {3},
issn = {0014-4754},
address = {Cham (ZG)},
publisher = {Springer International Publishing AG},
reportid = {PUBDB-2022-05338},
pages = {138},
year = {2022},
abstract = {Peptidyl-prolyl isomerases (PPIases) catalyze intrinsically
slow and often rate-limiting isomerization of prolyl-peptide
bonds in unfolded or partially folded proteins, thereby
speeding up the folding process and preventing misfolding.
They often possess binding and chaperone domains in addition
to the domain carrying the isomerization activity. Although
generally, their substrates display no identity in their
amino acid sequence upstream and downstream of the proline
with 20 possibilities for each residue, PPIases are
efficient enzymes. SlyD is a highly efficient PPIase
consisting of an isomerase domain and an additional
chaperone domain. The binding of peptide substrates to SlyD
and its enzymatic activity depend to some extend on the
proline-proximal residues, however, the impact of
proline-distant residues has not been investigated so far.
Here, we introduce a label-free NMR-based method to measure
SlyD activity on different peptide substrates and analysed
the data in the context of obtained binding affinities and
several co-crystal structures. We show that especially
charged and aromatic residues up to eight positions
downstream and three positions upstream of the proline and
outside the canonical region of similar conformations affect
the activity and binding, although they rarely display
distinct conformations in our crystal structures. We
hypothesize that these positions primarily influence the
association reaction. In the absence of the chaperone domain
the isomerase activity strongly correlates with substrate
affinity, whereas additional factors play a role in its
presence. The mutual orientation of isomerase and chaperone
domains depends on the presence of substrates in both
binding sites, implying allosteric regulation of enzymatic
activity.},
cin = {EMBL / CSSB-EMBL / EMBL-User / CSSB-EMBL-CL},
ddc = {610},
cid = {I:(DE-H253)EMBL-20120731 / I:(DE-H253)CSSB-EMBL-20141216 /
I:(DE-H253)EMBL-User-20120814 /
I:(DE-H253)CSSB-EMBL-CL-20210806},
pnm = {6G3 - PETRA III (DESY) (POF4-6G3)},
pid = {G:(DE-HGF)POF4-6G3},
experiment = {EXP:(DE-H253)P-P13-20150101 / EXP:(DE-H253)P-P14-20150101},
typ = {PUB:(DE-HGF)16},
pubmed = {35184231},
UT = {WOS:000757934900001},
doi = {10.1007/s00018-022-04179-4},
url = {https://bib-pubdb1.desy.de/record/483482},
}