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@ARTICLE{Blanke:418371,
author = {Blanke, Alexander and Pinheiro, M. and Watson, P. J. and
Fagan, M. J.},
title = {{A} biomechanical analysis of prognathous and orthognathous
insect head capsules: evidence for a many-to-one mapping of
form to function},
journal = {Journal of evolutionary biology},
volume = {31},
number = {5},
issn = {1010-061X},
address = {Oxford [u.a.]},
publisher = {Wiley-Blackwell},
reportid = {PUBDB-2019-00403},
pages = {665 - 674},
year = {2018},
note = {© European Society For Evolutionary Biology. Journal of
Evolutionary Biology © European Society For Evolutionary
Biology; Post referee fulltext in progress 2; Embargo 12
months from publication},
abstract = {Insect head shapes are remarkably variable, but the
influences of these changes on biomechanical performance are
unclear. Among ‘basal’ winged insects, such as
dragonflies, mayflies, earwigs and stoneflies, some of the
most prominent anatomical changes are the general mouthpart
orientation, eye size and the connection of the endoskeleton
to the head. Here, we assess these variations as well as
differing ridge and sclerite configurations using modern
engineering methods including multibody dynamics modelling
and finite element analysis in order to quantify and compare
the influence of anatomical changes on strain in particular
head regions and the whole head. We show that a range of
peculiar structures such as the genal/subgenal, epistomal
and circumocular areas are consistently highly loaded in all
species, despite drastically differing morphologies in
species with forward‐projecting (prognathous) and
downward‐projecting (orthognathous) mouthparts.
Sensitivity analyses show that the presence of eyes has a
negligible influence on head capsule strain if a
circumocular ridge is present. In contrast, the connection
of the dorsal endoskeletal arms to the head capsule
especially affects overall head loading in species with
downward‐projecting mouthparts. Analysis of the relative
strains between species for each head region reveals that
concerted changes in head substructures such as the subgenal
area, the endoskeleton and the epistomal area lead to a
consistent relative loading for the whole head capsule and
vulnerable structures such as the eyes. It appears that
biting‐chewing loads are managed by a system of
strengthening ridges on the head capsule irrespective of the
general mouthpart and head orientation. Concerted changes in
ridge and endoskeleton configuration might allow for more
radical anatomical changes such as the general mouthpart
orientation, which could be an explanation for the
variability of this trait among insects. In an evolutionary
context, many‐to‐one mapping of strain patterns onto a
relatively similar overall head loading indeed could have
fostered the dynamic diversification processes seen in
insects.},
cin = {DOOR},
ddc = {570},
cid = {I:(DE-H253)HAS-User-20120731},
pnm = {6G3 - PETRA III (POF3-622) / FS-Proposal: I-20120065
(I-20120065)},
pid = {G:(DE-HGF)POF3-6G3 / G:(DE-H253)I-20120065},
experiment = {EXP:(DE-H253)P-P05-20150101 / EXP:(DE-H253)D-BW2-20150101},
typ = {PUB:(DE-HGF)16},
pubmed = {pmid:29444377},
UT = {WOS:000432011600004},
doi = {10.1111/jeb.13251},
url = {https://bib-pubdb1.desy.de/record/418371},
}