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@ARTICLE{Seydel:390810,
author = {Seydel, Ulrich and Koch, Michel H. J. and Brandenburg,
Klaus},
title = {{S}tructural {P}olymorphisms of {R}ough {M}utant
{L}ipopolysaccharides {R}d to {R}a from {S}almonella
minnesota},
journal = {Journal of structural biology},
volume = {110},
number = {3},
issn = {1047-8477},
address = {San Diego, Calif.},
publisher = {Elsevier},
reportid = {PUBDB-2017-09865},
pages = {232 - 243},
year = {1993},
note = {EMBL},
abstract = {The structural polymorphisms of rough mutant
lipopolysaccharides (LPS) Rd, Rc, Rb, and Ra from Salmonella
minnesota (strains R4, R7, Rz, R5, R345, and R60,
respectively) were investigated as a function of
temperature, water content, and Mg$^{2+}$ concentration. The
gel to liquid crystalline (B↔α) phase transition
temperature (Tc) and the state of order within each phase
were measured by Fourier transform infrared spectroscopy.
The amount of bound water was determined by differential
scanning calorimetry and the three-dimensional structures in
each phase state were characterized by synchrotron radiation
X-ray diffraction. The results indicate an extremely complex
dependence of the structural behavior of LPS on ambient
conditions. The B↔α acyl chain melting temperatures at
high water contents (95-97\%), Tc = 31 to 32°C for LPS Rd,
33 to 35°C for LPS Rc to Rb, and 36°C for LPS Ra, increase
with decreasing water content and in the presence of
Mg$^{2+}$ cations with a concomitant broadening of the
transition range. Below 30 to 50\% water content, no
distinct phase transitions can be observed. These effects
are most pronounced for LPS with the shortest sugar chains.
Below 50\% water content, only lamellur structures can be
observed in the temperature range 5 to 80°C, independent of
the Mg$^{2+}$ concentration. Above 50\% water concentration,
for large [LPS]:[Mg$^{2+}$] molar ratios the predominant
structure above Tc is nonlamellar; for smaller
[LPS]:[Mg$^{2+}$] molar ratios a superposition of lamellar
and nonlamellar structures is found. For all LPS Rd to Rb at
low Mg$^{2+}$ concentrations, the phase transition is
connected with a change in the three-dimensional structure
from lamellar or mixed lamellar/nonlamellar to a pure
nonlamellar, probably cubic structure. The tendency to form
nonlamellar structures decreases with the length of the core
oligosaccharide. At an equimolar ratio of [LPS] and
[Mg$^{2+}$] a multibilayered organization is observed. Some
of the nonlamellar structures are cubic phases with
periodicities between 12 and 16 nm. The molecular dimensions
of the single endotoxin molecules in the absence and the
presence of external water are estimated from the different
lamellar periodicities, including those of free lipid A and
deep rough mutant LPS Re. These observations are discussed
with respect to the biological importance of LPS as a potent
inducer of biological effects in mammals.},
cin = {DESY(-2012)},
ddc = {540},
cid = {$I:(DE-H253)DESY_-2012_-20170516$},
pnm = {899 - ohne Topic (POF3-899)},
pid = {G:(DE-HGF)POF3-899},
experiment = {EXP:(DE-MLZ)NOSPEC-20140101},
typ = {PUB:(DE-HGF)16},
UT = {WOS:A1993LU06900008},
pubmed = {pmid:8373704},
doi = {10.1006/jsbi.1993.1026},
url = {https://bib-pubdb1.desy.de/record/390810},
}