TY - JOUR
AU - Mastny, Markus
AU - Heuck, Alexander
AU - Kurzbauer, Robert
AU - Heiduk, Anja
AU - Boisguerin, Prisca
AU - Volkmer, Rudolf
AU - Ehrmann, Michael
AU - Rodrigues, Christopher D. A.
AU - Rudner, David Z.
AU - Clausen, Tim
TI - CtpB Assembles a Gated Protease Tunnel Regulating Cell-Cell Signaling during Spore Formation in Bacillus subtilis
JO - Cell
VL - 155
IS - 3
SN - 0092-8674
CY - [Cambridge, Mass.]
PB - Cell Press
M1 - DESY-2014-02420
SP - 647 - 658
PY - 2013
N1 - (c) Elsevier Inc.
AB - Spore formation in Bacillus subtilis relies on a regulated intramembrane proteolysis (RIP) pathway that synchronizes mother-cell and forespore development. To address the molecular basis of this SpoIV transmembrane signaling, we carried out a structure-function analysis of the activating protease CtpB. Crystal structures reflecting distinct functional states show that CtpB constitutes a ring-like protein scaffold penetrated by two narrow tunnels. Access to the proteolytic sites sequestered within these tunnels is controlled by PDZ domains that rearrange upon substrate binding. Accordingly, CtpB resembles a minimal version of a self-compartmentalizing protease regulated by a unique allosteric mechanism. Moreover, biochemical analysis of the PDZ-gated channel combined with sporulation assays reveal that activation of the SpoIV RIP pathway is induced by the concerted activity of CtpB and a second signaling protease, SpoIVB. This proteolytic mechanism is of broad relevance for cell-cell communication, illustrating how distinct signaling pathways can be integrated into a single RIP module.
LB - PUB:(DE-HGF)16
UR - <Go to ISI:>//WOS:000326571800018
C6 - pmid:24243021
DO - DOI:10.1016/j.cell.2013.09.050
UR - https://bib-pubdb1.desy.de/record/168215
ER -