% IMPORTANT: The following is UTF-8 encoded. This means that in the presence
% of non-ASCII characters, it will not work with BibTeX 0.99 or older.
% Instead, you should use an up-to-date BibTeX implementation like “bibtex8” or
% “biber”.
@PHDTHESIS{Chretien:626020,
author = {Chretien, Anais},
othercontributors = {Pearson, Arwen and Meier, Chris},
title = {{E}lucidating signal transduction in multi-domain {BLUF}
photoreceptors by studying the {P}hotoactivated {A}denylate
{C}yclase {O}a{PAC}},
school = {University of Hamburg},
type = {Dissertation},
reportid = {PUBDB-2025-01257},
pages = {258},
year = {2024},
note = {Dissertation, University of Hamburg, 2024},
abstract = {Photosensory receptors, essential molecular entities across
all domains of life, enable organisms to detect and respond
to light stimuli, underpinning their critical involvement in
regulating biological processes such as phototropism,
circadian rhythms, photomorphogenesis, and photosynthesis.
Among the myriad types of photosensory receptors, blue light
sensing proteins such as Blue Light Using Flavin (BLUF)
photoreceptors distinguish themselves through their ability
to utilize blue light for signalling. Characterized by the
conserved structure of their sensor domain, BLUF
photoreceptors are found in a wide array of organisms, from
bacteria and algae to plants and certain fungi. Known for
their capacity to bind flavin chromophores, typically flavin
adenine dinucleotide (FAD), they undergo conformational
changes upon blue photon absorption, leading to downstream
signalling events, highlighting their pivotal role in the
adaptive responses of various organisms to light. This
dissertation provides a comprehensive exploration of the
BLUF photoreceptors, particularly focusing on the
Photoactivated Adenylate Cyclase protein from Oscillatoria
acuminata (OaPAC), which comprises a BLUF sensor domain
linked to an Adenylate Cyclase (AC) effector domain,
catalysing the conversion of ATP into cAMP. This study aims
to elucidate the photoactivation mechanism of OaPAC and the
ensuing signal transduction pathway, employing an
integrative approach that leverages time-resolved
crystallography, small angle X-ray scattering, spectroscopy,
and biochemical characterization techniques. Special
emphasis is placed on the Tyr-Gln-Met triad in the BLUF
domain, which plays a crucial role in the initial
light-induced rearrangements. Additionally, significant
attention is given to the less understood aspects of BLUF
photoreceptors, particularly the transduction of the initial
light signal to more distal parts of the protein, which
ultimately leads to biological activity. This research
identifies a Met(out)/Trp(in) transition as a crucial
element in conveying the signal to the α-helix linker
region. Finally, structural models of OaPAC with ATP bound
in the active site, along with complementary FTIR
investigations, provide a thorough understanding of ATP
binding and allosteric communication. As a result, the
research presented in this dissertation not only expands the
fundamental understanding of BLUF photoreceptor biology, but
also provides a framework for future studies aimed at
deciphering complete signal transduction pathways in
multi-domain BLUF photoreceptors and towards the development
of optogenetic tools.},
pnm = {PHGS, VH-GS-500 - PIER Helmholtz Graduate School
$(2015_IFV-VH-GS-500)$},
pid = {$G:(DE-HGF)2015_IFV-VH-GS-500$},
typ = {PUB:(DE-HGF)11},
urn = {urn:nbn:de:gbv:18-ediss-121292},
url = {https://bib-pubdb1.desy.de/record/626020},
}